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<jats:title>Functional ocean biogeography</jats:title> <jats:p> Marine ecosystems are well represented in metagenomic and transcriptomic data. These data are not routinely used to test ecosystem models that explore ocean biogeography or biogeochemistry. Coles <jats:italic>et al.</jats:italic> built a model in which genes for a range of functions were assigned to different suites of simulated microbes (see the Perspective by Rynearson). Communities emerged from the model with realistic biogeographical and biogeochemical profiles when compared to microbial data collected from the Amazon River plume. However, functional composition trumped the details of taxonomy, and different, coevolving community compositions emerged that provided similar biogeochemical outcomes. </jats:p> <jats:p> <jats:italic>Science</jats:italic> , this issue p. <jats:related-article xmlns:xlink="http://www.w3.org/1999/xlink" ext-link-type="doi" issue="6367" page="1149" related-article-type="in-this-issue" vol="358" xlink:href="10.1126/science.aan5712">1149</jats:related-article> ; see also p. <jats:related-article xmlns:xlink="http://www.w3.org/1999/xlink" ext-link-type="doi" issue="6367" page="1129" related-article-type="in-this-issue" vol="358" xlink:href="10.1126/science.aar3431">1129</jats:related-article> </jats:p>