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<jats:p> A video introduction to the year's top science story, featuring scientists C. Owen Lovejoy, Tim White, Giday WoldeGabriel, Yohannes Haile-Selassie, <jats:italic>Science</jats:italic> contributing correspondent Ann Gibbons, and commentary by paleoanthropologist Andrew Hill. </jats:p>

<jats:p>A rare 4.4-million-year-old skeleton has drawn back the curtain of time to reveal the surprising body plan and ecology of our earliest ancestors.</jats:p>

<jats:p>This year's runners-up for Breakthrough of the Year include new gamma-ray observations, the long-sought receptor for a key plant hormone, mock monopoles, a drug that increases life span, ice on the moon, gene-therapy successes, insights into the properties of graphene and how to use it to make novel devices, Hubble's rebirth, and the first x-ray laser.</jats:p>

<jats:p> <jats:italic>Science</jats:italic> 's editors clearly foresaw this year's burgeoning of plant genome sequences, progress on emissions reductions in the run-up to the U.N. conference in Copenhagen, and the continued failure to spot dark matter. Last year's other predictions will take more time to come to fruition. </jats:p>

<jats:p>The global financial crisis made 2009 a tough year for many U.S. academic institutions dependent on state funding or endowments. But it has also been a banner year for thousands of individual scientists, whose labs have benefited from billions of dollars in U.S. government stimulus funding.</jats:p>

<jats:p> In 2010, <jats:italic>Science</jats:italic> 's editors will be watching for developments with induced pluripotent stem cells, the Alpha Magnetic Spectrometer, "exome sequencing," disrupting the metabolism of tumor cells, and human space flight. </jats:p>